Mega Man misterious wakes up one day in a familiar place. He's on the old destroyed highway that he traveled on way back in Mega Man X1. At the end of the highway he meets up with Vile. After fighting Vile, X is transported back to the Dr. Cain's lab. The computer technician, Middy, explains to X that someone has reprogrammed the main frame computer and has brought old battle data back to life. Selflessly, X allows himself to be digitally projected into the computer to fight the menace (As usual, Zero stays behind to "Fight the mavericks").
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